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Testudo quadratus Cope 1884 (gopher tortoise)

Reptilia - Testudines - Testudinidae

PaleoDB taxon number: 105215

Alternative combination: Geochelone quadrata

Full reference: E. D. Cope. 1884. Testudinata. The Vertebrata of the Tertiary formations of the West. In: Report of the United States Geological Survey of the Territories 3:111-150

Belongs to Testudinidae according to E. Vlachos 2018

Sister taxa: Achilemys, Agrionemys, Aldabrachelys, Anhuichelys, Astrochelys, Cachuga, Caryoderma, Centrochelys, Cheirogaster, Chelone, Chelonoidis, Chelys patagonica, Chersina, Cylindraspis, Damonia, Dipsochelys, Emys sombrerensis, Ergilemys, Eurotestudo, Floridemys, Floridemys hurdi, Fontainechelon, Geochelone, Geochelone (Chelonoidis), Geochelone (Hesperotestudo) equicomes, Gigantochersina, Gopherus, Gopherus praecedens, Hesperotestudo, Homopus, Impregnochelys, Indotestudo, Juvemys, Kansuchelys, Kinixys, Malacochersus, Manouria, Megalochelys, Mesochersus, Miotestudo, Namibchersus, Nicoria, Palaeoemys, Paleotestudo, Pelorochelon, Psammobates, Pyxis, Sinohadrianus, Stigmochelys, Stylemys, Stylemys calaverensis, Taraschelon, Testudininae, Testudininei, Testudinites sellovii, Testudo, Testudo annae, Testudo atascosae, Testudo castrensis, Testudo cultrata, Testudo exornata, Testudo formosa, Testudo francisi, Testudo hayi, Testudo klettiana, Testudo laticaudata, Testudo louisekressmani, Testudo luciae, Testudo ocalana, Testudo paranensis, Testudo peragrans, Testudo undata, Titanochelon

Type specimen: AMNH 1149 (holotype), a partial shell (fragmentary epiplastron and partial nuchal). Its type locality is Horsetail Creek, which is in a Chadronian terrestrial horizon in the White River Formation of Colorado.

Ecology: ground dwelling herbivore-herbivore