PaleoDB taxon number: 199232
Full reference: A. V. Martynov. 1937. Permian fossil insects from Kargala and their relationships. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7:1-92
Parent taxon: Coleoptera according to A. V. Martynov 1937
Sister taxa: Acmaeodera perreaui, Adephaga, Adikia, Adynasia, Allognosis, Anancaeon, Anapiptus, Ancylocheira liasina, Angelinella, Anhuistoma, Anhydrophilus, Anomerus, Anypostatus, Apheloodes, Apiaria, Apicasia, Apioderes, Apiopyrenides, Apistotes, Apsychus, Aptilotitus, Archostemata, Archosyne, Auchenophorites, Bareus, Bathygerus, Bellingera, Bellingeropsis, Blaptoides, Bostrichini, Bothroptera, Brachylaimon, Brachytrachelites, Brevispinnatus, Brodiola, Bruchelae, Bucklandula, Buprestium, Carabicina, Carabiques, Carabocera, Cardioides, Cebrionates, Cerambicini, Cerylonopsis, Chimaerocoleites, Chlaeniopsis, Chrysomelidium, Chrysomelophana, Cistela, Cistelites, Clerii, Clinomerus, Coccinellophana, Coleopteron, Coleopteron depressus, Coleopteron interpunctatus, Coleopteron laevigatus, Coleopteron scabriusculus, Colymbetopsis, Coprophagi, Cretaceites, Cretofalselaterium, Cretogermen, Criocerina pristina, Critoderma, Cryptauchenia, Cucujipes, Curculidium, Curculionites, Curculionites liasinus, Curculium, Cydnopsis nigromembranacea, Cyphospheron, Darbiopelta, Diachoristes, Diaperialae, Diaperidium, Diatarastus, Diatrypamene, Dicyphelus, Diphymation, Diplocelides, Diplothece, Doggeria, Dysarestus, Ecthlimma, Elateridopsidae, Elateridopsis, Elytrulum, Enamma, Entomocantharus, Entomophagi, Eocoleopteron, Episcepes, Epomenus, Erotilenae, Erotylites, Etheridgea, Euchroma arvernica, Eurynotellus, Eurysphinctus, Eusarcantarus, Explansimarginis, Flichea, Gastrodelus, Gastroratus, Geotrupini, Glaphoptera, Glaphyropterites, Glaphyropterula, Grahamelytron, Grasselites, Gurvanocoleus, Gyrinites, Gyrinulopsis, Halticophana, Harpalidium, Heeriaopsis, Helophoropsis, Helophorus exilis, Helophorus magnus, Helopidium, Helopii, Helopium, Hubeicoleus, Hydrobius godeti, Hydrobius obsoletus, Hydrocanthares, Hydroicetes, Hydrophilites, Hydrophilus stenopterus, Hydroporopsis, Hydrous ebeninus, Hylesinus lineatus, Hyperomima, Ipidae, Jurodidae, Kamaroma, Katapiptus, Keleusticus, Kibdelia, Kryptophagus, Laimocenos, Lamiites, Lamiophanes, Laodiscis, Latridiites, Leioodes, Leptomites, Leptosolenophorus, Longimaxilla, Loxocamarotus, Macrogastri, Macrotrachelites, Malacodermi, Malacodermidae, Masselytron, Megacentrus, Megachorites, Melanophilites, Melanophilopsis, Mesoncus, Mesorhynchophora, Mesotricupes, Mesotylites, Metanastes, Microcoleopteron, Microentomus, Mordellonae, Myxophaga, Nannoodes, Nebrioides, Necrophagi, Necticus minutus, Nitidulariae, Nubili, Omogongylus, Onkedodimus, Onthophagus luteus, Ooidellus, Ooperiglyptus, Ooperioristus, Oropseini, Oxycephalites, Oxytoroptera, Pachycoleon, Pachypleurites, Paklapwania, Palaeotrachys, Palaeotylinae, Pallax, Palpatores, Pantodapus, Paracurculium, Paragyrinus, Parasilphites, Parnidium, Parnosoma, Pedinosoma, Peltocoleops, Periboloptera, Peridosoma, Permocoleus, Petalocupes, Petrorophus, Phaulogyrinus, Pholipheron, Phytoplesion, Pimeliodes, Plastobuprestites, Plastonebria, Platyprosopi, Pleuralocista, Podabrus santaritensis, Polypamon, Polyphaga, Ponomarenkiidae, Prionophana, Prochrysomela, Proheuristes, Prophasis, Prosthecogmus, Prosthenostictus, Protocoleoptera, Protocuneus, Pselaphii, Pseudobuprestites, Pseudocarabites, Pseudocurculionites, Pseudocymindis, Pseudocyphon, Pseudoelateropsis, Pseudohydrophilites, Pseudoprionites, Pseudotelephorus, Pseudotenebrio, Pseudus, Ptiniores, Pyrochroophana, Rhomaleus, Rhynchophori, Saperda valdensis, Saprophagi, Scalopoides, Scarabaeides, Semiglobus, Sideriosemion, Silphidium, Sinoelaterium, Sitonites, Smodicoptera, Sojanocoleidae, Sphaericites, Sphaeridiota, Staphylini, Sternoxi, Stictulus, Strongylites, Strullendorfia, Telephorium, Tenebrionites, Tetillopsis, Tetragonides, Thurmannia, Toksunius, Tolype, Triadocoleopteron, Trichelepturgetes, Tridigitati, Tripsalis, Trixagites, Trochiscites, Trochmalus, Trogossitarii, Trypocoleus, Tychon, Ulomites, Velenovskya, Versicolores, Wollastonia, Xenogyrinus, Xylophagi, Xylotupia, Yanqingia, Zetemenos, Tachypus
Subtaxa: none
Ecology: