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†suborder Protocoleoptera Tillyard 1924 (beetle)

Insecta - Coleoptera

PaleoDB taxon number: 212840

Full reference: R. J. Tillyard. 1924. Upper Permian Coleoptera and a new order from the Belmont Beds, New South Wales. Proceedings of the Linnean Society of New South Wales 49:429-435

Parent taxon: Coleoptera according to P. Bouchard et al. 2011

Sister taxa: Acmaeodera perreaui, Adephaga, Adikia, Adynasia, Allognosis, Anancaeon, Anapiptus, Ancylocheira liasina, Angelinella, Anhuistoma, Anhydrophilus, Anomerus, Anypostatus, Apheloodes, Apiaria, Apicasia, Apioderes, Apiopyrenides, Apistotes, Apsychus, Aptilotitus, Archostemata, Archosyne, Auchenophorites, Bareus, Bathygerus, Bellingera, Bellingeropsis, Blaptoides, Bostrichini, Bothroptera, Bothynophora, Brachylaimon, Brachytrachelites, Brevispinnatus, Brodiola, Bruchelae, Bucklandula, Buprestium, Camptodontus, Carabicina, Carabiques, Carabocera, Cardioides, Caryoborus, Cebrionates, Cerambicini, Cerylonopsis, Chimaerocoleites, Chlaeniopsis, Chrysomelidium, Chrysomelophana, Cistela, Clerii, Clinomerus, Coccinellophana, Coleopteron, Coleopteron depressus, Coleopteron interpunctatus, Coleopteron laevigatus, Coleopteron scabriusculus, Colymbetopsis, Coprophagi, Cretofalselaterium, Cretogermen, Criocerina pristina, Critoderma, Cryptauchenia, Cucujipes, Curculidium, Curculionites, Curculionites liasinus, Curculiopsidae, Curculium, Cydnopsis nigromembranacea, Cyphospheron, Darbiopelta, Diachoristes, Diaperialae, Diaperidium, Diatarastus, Diatrypamene, Dicyphelus, Diphymation, Diplocelides, Diplothece, Doggeria, Doggeriopsis, Ecthlimma, Elateridopsidae, Elateridopsis, Elytrulum, Enamma, Entomocantharus, Entomophagi, Eocoleopteron, Episcepes, Epomenus, Erotilenae, Erotylites, Etheridgea, Euchroma arvernica, Eurynotellus, Eurysphinctus, Eusarcantarus, Explansimarginis, Flichea, Gastrodelus, Gastroratus, Geotrupini, Glaphoptera, Glaphyropterites, Glaphyropterula, Grahamelytron, Grasselites, Gurvanocoleus, Gyrinites, Gyrinulopsis, Halticophana, Harpalidium, Heeriaopsis, Helophoropsis, Helophorus exilis, Helophorus magnus, Helopides, Helopidium, Helopii, Helopium, Holcoptera, Hubeicoleus, Hydrobius godeti, Hydrobius obsoletus, Hydrocanthares, Hydroicetes, Hydrophilites, Hydrophilus stenopterus, Hydroporopsis, Hydrous ebeninus, Hylesinus lineatus, Hyperomima, Ipidae, Ironicus, Jurodidae, Kamaroma, Katapiptus, Katapontisus, Keleusticus, Kelidus, Kibdelia, Kryptophagus, Laimocenos, Lamiites, Lamiophanes, Latridiites, Leioodes, Leptomites, Leptosolenophorus, Longimaxilla, Loxocamarotus, Macrogastri, Macrotrachelites, Malacodermi, Malacodermidae, Masselytron, Megacentrus, Megachorites, Melanophilites, Melanophilopsis, Mesoncus, Mesorhynchophora, Mesosagrites, Mesotylites, Metanastes, Microcoleopteron, Mordellonae, Myxophaga, Nannocurculionites, Nannoodes, Nebrioides, Necrophagi, Necticus minutus, Nitidulariae, Nubili, Omogongylus, Onkedodimus, Onthophagus luteus, Ooidellus, Ooperiglyptus, Ooperioristus, Oxycephalites, Oxytoroptera, Pachycoleon, Pachypleurites, Paklapwania, Palaeotrachys, Pallax, Palpatores, Pantodapus, Parabuprestites, Paracurculionites, Paracurculium, Paradoggeria, Paragyrinus, Parasilphites, Parnidium, Parnosoma, Pedinosoma, Peltocoleops, Periboloptera, Peridosoma, Permocoleus, Petrorophus, Phanerogramma, Phaulogyrinus, Pholipheron, Phytoplesion, Pimeliodes, Plastobuprestites, Plastonebria, Platyprosopi, Pleuralocista, Polypamon, Polyphaga, Prionophana, Prochrysomela, Proheuristes, Prophasis, Prosthecogmus, Prosthenostictus, Protocuneus, Pselaphii, Pseudobuprestites, Pseudocarabites, Pseudocurculionites, Pseudocymindis, Pseudocyphon, Pseudoelateropsis, Pseudohydrophilites, Pseudomesalia, Pseudoprionites, Pseudosilphites, Pseudotelephorus, Pseudotenebrio, Pseudus, Ptiniores, Pyrochroophana, Rhomaleus, Rhynchophori, Saperda valdensis, Saprophagi, Scalopoides, Scarabaeides, Semiglobus, Sideriosemion, Silphidium, Sinoelaterium, Sitonites, Smodicoptera, Sojanocoleidae, Sphaericites, Sphaeridiota, Staphylini, Sternoxi, Stictulus, Stigmenamma, Strongylites, Strullendorfia, Telephorium, Tenebrionites, Tentyridium, Tetillopsis, Tetragonides, Thurmannia, Toksunius, Tolype, Triadocoleopteron, Trichelepturgetes, Tridigitati, Tripsalis, Trixagites, Trochiscites, Trochmalus, Trogossitarii, Trypocoleus, Tychon, Ulomites, Velenovskya, Versicolores, Wollastonia, Xenogyrinus, Xylophagi, Xylotupia, Yanqingia, Zetemenos

Subtaxa: Permocupedoidea, Permosynoidea, Tshekardocoleoidea

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