PaleoDB taxon number: 229485
Full reference: H. Howard. 1958. Miocene sulids of southern California. Contributions in Science 25:1-15
Sister taxa: Sula (Sula), Sula arvernensis, Sula brandi, Sula clarki, Sula dactylatra, Sula figueroae, Sula granti, Sula guano, Sula leucogaster, Sula magna, Sula nebouxii, Sula phosphata, Sula sula, Sula sulita, Sula universitatis, Sula variegata, Sula willetti
Type specimen: LACM 2674, a set of limb elements (right and left wing girdle). Its type locality is Ventura Boulevard & Whitsett Avenue, which is in a Miocene marine shale in California.
Ecology: ground dwelling carnivore