PaleoDB taxon number: 260620
Full reference: O. Heer. 1856. Ueber die fossilen Insekten von Aix in der Provence. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich 1:1-40
Belongs to Coleoptera according to M. Fikacek et al. 2010
Sister taxa: Acmaeodera perreaui, Adephaga, Adikia, Adynasia, Allognosis, Anancaeon, Anapiptus, Ancylocheira liasina, Angelinella, Anhuistoma, Anhydrophilus, Anomerus, Anypostatus, Apheloodes, Apiaria, Apicasia, Apioderes, Apiopyrenides, Apistotes, Apsychus, Aptilotitus, Archostemata, Archosyne, Auchenophorites, Bareus, Bathygerus, Bellingera, Bellingeropsis, Blaptoides, Bostrichini, Bothroptera, Bothynophora, Brachylaimon, Brachytrachelites, Brevispinnatus, Brodiola, Bruchelae, Bucklandula, Buprestium, Camptodontus, Carabicina, Carabiques, Carabocera, Cardioides, Caryoborus, Cebrionates, Cerambicini, Cerylonopsis, Chimaerocoleites, Chlaeniopsis, Chrysomelidium, Chrysomelophana, Cistela, Clerii, Clinomerus, Coccinellophana, Coleopteron, Coleopteron depressus, Coleopteron interpunctatus, Coleopteron laevigatus, Coleopteron scabriusculus, Colymbetopsis, Coprophagi, Cretofalselaterium, Cretogermen, Criocerina pristina, Critoderma, Cryptauchenia, Cucujipes, Curculidium, Curculionites, Curculionites liasinus, Curculiopsidae, Curculium, Cydnopsis nigromembranacea, Cyphospheron, Darbiopelta, Diachoristes, Diaperialae, Diaperidium, Diatarastus, Diatrypamene, Dicyphelus, Diphymation, Diplocelides, Diplothece, Doggeria, Doggeriopsis, Ecthlimma, Elateridopsidae, Elateridopsis, Elytrulum, Enamma, Entomocantharus, Entomophagi, Eocoleopteron, Episcepes, Epomenus, Erotilenae, Erotylites, Etheridgea, Euchroma arvernica, Eurynotellus, Eurysphinctus, Eusarcantarus, Explansimarginis, Flichea, Gastrodelus, Gastroratus, Geotrupini, Glaphoptera, Glaphyropterites, Glaphyropterula, Grahamelytron, Grasselites, Gurvanocoleus, Gyrinites, Gyrinulopsis, Halticophana, Harpalidium, Heeriaopsis, Helophoropsis, Helophorus exilis, Helophorus magnus, Helopides, Helopidium, Helopii, Helopium, Holcoptera, Hubeicoleus, Hydrobius godeti, Hydrocanthares, Hydroicetes, Hydrophilites, Hydrophilus stenopterus, Hydroporopsis, Hydrous ebeninus, Hylesinus lineatus, Hyperomima, Ipidae, Ironicus, Jurodidae, Kamaroma, Katapiptus, Katapontisus, Keleusticus, Kelidus, Kibdelia, Kryptophagus, Laimocenos, Lamiites, Lamiophanes, Latridiites, Leioodes, Leptomites, Leptosolenophorus, Longimaxilla, Loxocamarotus, Macrogastri, Macrotrachelites, Malacodermi, Malacodermidae, Masselytron, Megacentrus, Megachorites, Melanophilites, Melanophilopsis, Mesoncus, Mesorhynchophora, Mesosagrites, Mesotylites, Metanastes, Microcoleopteron, Mordellonae, Myxophaga, Nannocurculionites, Nannoodes, Nebrioides, Necrophagi, Necticus minutus, Nitidulariae, Nubili, Omogongylus, Onkedodimus, Onthophagus luteus, Ooidellus, Ooperiglyptus, Ooperioristus, Oxycephalites, Oxytoroptera, Pachycoleon, Pachypleurites, Paklapwania, Palaeotrachys, Pallax, Palpatores, Pantodapus, Parabuprestites, Paracurculionites, Paracurculium, Paradoggeria, Paragyrinus, Parasilphites, Parnidium, Parnosoma, Pedinosoma, Peltocoleops, Periboloptera, Peridosoma, Permocoleus, Petrorophus, Phanerogramma, Phaulogyrinus, Pholipheron, Phytoplesion, Pimeliodes, Plastobuprestites, Plastonebria, Platyprosopi, Pleuralocista, Polypamon, Polyphaga, Prionophana, Prochrysomela, Proheuristes, Prophasis, Prosthecogmus, Prosthenostictus, Protocoleoptera, Protocuneus, Pselaphii, Pseudobuprestites, Pseudocarabites, Pseudocurculionites, Pseudocymindis, Pseudocyphon, Pseudoelateropsis, Pseudohydrophilites, Pseudomesalia, Pseudoprionites, Pseudosilphites, Pseudotelephorus, Pseudotenebrio, Pseudus, Ptiniores, Pyrochroophana, Rhomaleus, Rhynchophori, Saperda valdensis, Saprophagi, Scalopoides, Scarabaeides, Semiglobus, Sideriosemion, Silphidium, Sinoelaterium, Sitonites, Smodicoptera, Sojanocoleidae, Sphaericites, Sphaeridiota, Staphylini, Sternoxi, Stictulus, Stigmenamma, Strongylites, Strullendorfia, Telephorium, Tenebrionites, Tentyridium, Tetillopsis, Tetragonides, Thurmannia, Toksunius, Tolype, Triadocoleopteron, Trichelepturgetes, Tridigitati, Tripsalis, Trixagites, Trochiscites, Trochmalus, Trogossitarii, Trypocoleus, Tychon, Ulomites, Velenovskya, Versicolores, Wollastonia, Xenogyrinus, Xylophagi, Xylotupia, Yanqingia, Zetemenos
Type specimen: NHM P(IV) 39859, an exoskeleton. Its type locality is Aix-en-Provence (Murchison collection), which is in a Chattian lacustrine - large limestone in the Niveau du gypse d'Aix Formation of France.
Ecology:
Average measurements (in mm): body 7.70 x 4.70, elytron length 4.10