PaleoDB taxon number: 260620
Full reference: O. Heer. 1856. Ueber die fossilen Insekten von Aix in der Provence. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich 1:1-40
Belongs to Coleoptera according to M. Fiká?ek et al. 2010
Sister taxa: Acmaeodera perreaui, Adelidium, Adephaga, Adikia, Adynasia, Allognosis, Allopliosilpha, Anancaeon, Anapiptus, Ancylocheira liasina, Anepismus, Angelinella, Anhuistoma, Anhydrophilus, Anomerus, Anypostatus, Apheloodes, Apiaria, Apicasia, Apioderes, Apiopyrenides, Apistotes, Apsychus, Aptilotitus, Archostemata, Archosyne, Auchenophorites, Bareus, Bathygerus, Bellingera, Bellingeropsis, Blaptoides, Bostrichini, Bothroptera, Brachylaimon, Brachytrachelites, Brevispinnatus, Brochocoleus indibili, Brodiola, Bruchelae, Bucklandula, Buprestium, Carabicina, Carabiques, Carabocera, Cardioides, Cebrionates, Cerambicini, Cerylonopsis, Chimaerocoleites, Chlaeniopsis, Chrysomelidium, Chrysomelophana, Cistela, Cistelites, Clathropenna, Clerii, Clinomerus, Coccinellophana, Coleopteron, Coleopteron depressus, Coleopteron interpunctatus, Coleopteron laevigatus, Coleopteron scabriusculus, Colymbetopsis, Coprophagi, Cretaceites, Cretofalselaterium, Cretogermen, Criocerina pristina, Critoderma, Cryptauchenia, Cucujipes, Curculidium, Curculionites, Curculionites liasinus, Curculiopsidae, Curculium, Cydnopsis nigromembranacea, Cyphospheron, Darbiopelta, Diachoristes, Diaperialae, Diaperidium, Diatarastus, Diatrypamene, Dicyphelus, Diphymation, Diplocelides, Diplothece, Doggeria, Dysarestus, Ecthlimma, Elateridopsidae, Elateridopsis, Elytrulum, Enamma, Entomocantharus, Entomophagi, Eocoleopteron, Episcepes, Epomenus, Erotilenae, Erotylites, Etheridgea, Euchroma arvernica, Eurynotellus, Eurysphinctus, Eusarcantarus, Explansimarginis, Flichea, Fuscicupes, Gastrodelus, Gastroratus, Geotrupini, Glaphoptera, Glaphyropterites, Glaphyropterula, Grahamelytron, Grasselites, Gurvanocoleus, Gyrinites, Gyrinulopsis, Halticophana, Harpalidium, Heeriaopsis, Helophoropsis, Helophorus exilis, Helophorus magnus, Helopidium, Helopii, Helopium, Helops atticus, Hubeicoleus, Hydrobius godeti, Hydrocanthares, Hydroicetes, Hydrophilites, Hydrophilus stenopterus, Hydroporopsis, Hydrous ebeninus, Hylesinus lineatus, Hyperomima, Ipidae, Jurodidae, Kamaroma, Katapiptus, Keleusticus, Kibdelia, Kryptophagus, Laimocenos, Lamiites, Lamiophanes, Laodiscis, Latridiites, Leioodes, Leptomites, Leptosolenophorus, Longimaxilla, Loxocamarotus, Macrogastri, Macrotrachelites, Malacodermi, Malacodermidae, Masselytron, Megacentrus, Megachorites, Melanophilites, Melanophilopsis, Menephiloides, Mesoncus, Mesorhynchophora, Mesotricupes, Mesotylites, Metanastes, Microcoleopteron, Microentomus, Mordellonae, Myxophaga, Nannoodes, Nebrioides, Necrophagi, Necticus minutus, Nitidulariae, Notocupes multituberatus, Notocupes undatabdominus, Nubili, Omogongylus, Onkedodimus, Onthophagus luteus, Ooidellus, Ooperiglyptus, Ooperioristus, Oropseini, Oxycephalites, Oxytoroptera, Pachycoleon, Pachypleurites, Paklapwania, Palaeotrachys, Palaeotylinae, Pallax, Palpatores, Pantodapus, Paracurculium, Paragyrinus, Parasilphites, Parnidium, Parnosoma, Pedinosoma, Peltocoleops, Periboloptera, Peridosoma, Petalocupes, Petrorophus, Phaulogyrinus, Pholipheron, Phytoplesion, Pimeliodes, Plastobuprestites, Plastonebria, Platyprosopi, Pleuralocista, Podabrus santaritensis, Polypamon, Polyphaga, Ponomarenkiidae, Prionophana, Prochrysomela, Proheuristes, Prophasis, Prosthecogmus, Prosthenostictus, Protocoleoptera, Protocuneus, Pselaphii, Pseudobuprestites, Pseudocarabites, Pseudocurculionites, Pseudocymindis, Pseudocyphon, Pseudoelateropsis, Pseudohydrophilites, Pseudoprionites, Pseudotelephorus, Pseudotenebrio, Pseudus, Ptiniores, Rhomaleus, Rhynchophori, Saperda valdensis, Saprophagi, Scalopoides, Scarabaeides, Semiglobus, Sideriosemion, Silphidium, Sinoelaterium, Sitonites, Smodicoptera, Sojanocoleidae, Sphaericites, Sphaeridiota, Staphylini, Sternoxi, Stictulus, Strongylites, Strullendorfia, Telephorium, Tenebrionites, Tetillopsis, Tetragonides, Thurmannia, Toksunius, Tolype, Triadocoleopteron, Trichelepturgetes, Tridigitati, Tripsalis, Trixagites, Trochiscites, Trochmalus, Trogossitarii, Trypocoleus, Tychon, Ulomites, Velenovskya, Versicolores, Wollastonia, Xenogyrinus, Xylophagi, Xylotupia, Yanqingia, Zetemenos, Tachypus
Type specimen: NHM P(IV) 39859, an exoskeleton. Its type locality is Aix-en-Provence (Murchison collection), which is in a Chattian lacustrine - large limestone in the Niveau du gypse d'Aix Formation of France.
Ecology:
Average measurements (in mm): body 7.70 x 4.70, elytron length 4.10