PaleoDB taxon number: 302959
Full reference: I. Fujiyama. 1968. A Miocene fossil of tropical dung beetle from Noto, Japan (Tertiary insect fauna of Japan, 2). Bulletin of the National Science Museum 11:201-210
Sister taxa: none
Type specimen: Kyoto University JC 500528, an exoskeleton. Its type locality is Takaya, Suzu-shi, Noto Peninsula, which is in a Miocene terrestrial siliciclastic in the Yanagida Formation of Japan.
Ecology: epifaunal grazer
Average measurements (in mm): elytron 28.5 x 16.0