PaleoDB taxon number: 358822
Full reference: E. A. Zvonok, N. V. Zelenkov, and I. G. Danilov. 2016. A new unusual waterbird (Aves, ?Suliformes) from the Eocene of Kazakhstan. Journal of Vertebrate Paleontology 36(1):e1035783
Sister taxa: none
Type specimen: PIN 2612/2, a limb element (proximal fragment of the left humerus). Its type locality is Kuyulus, which is in a Bartonian coastal horizon in the Shorym Formation of Kazakhstan.