PaleoDB taxon number: 379353
Full reference: J. X. Samuels, K. E. Bredehoeft, and S. C. Wallace. 2018. A new species of Gulo from the Early Pliocene Gray Fossil Site (Eastern United States); rethinking the evolution of wolverines. PeerJ 6:e4648
Type specimen: ETMNH 3663, a maxilla (partial right maxilla with P2 and P4). Its type locality is Gray Fossil Site, which is in a Hemphillian/Blancan sinkhole siltstone/conglomerate in Tennessee.