PaleoDB taxon number: 227464
Full reference: D. T. Ksepka, R. E. Fordyce, T. Ando and C. M. Jones. 2012. New fossil penguins (AVes, Sphenisciformes) from the Oligocene of New Zealand reveal the skeletal plan of stem penguins. Journal of Vertebrate Paleontology 32(2):235-254
Sister taxon: Kairuku grebneffi
Type specimen: OU 12652, a partial skeleton. Its type locality is J40/f190 - Waihao River [Kokoamu Greensand], which is in a Chattian offshore shelf sandstone in the Kokoamu Greensand Formation of New Zealand.
Ecology: ground dwelling carnivore