PaleoDB taxon number: 227466
Full reference: D. T. Ksepka, R. E. Fordyce, T. Ando and C. M. Jones. 2012. New fossil penguins (AVes, Sphenisciformes) from the Oligocene of New Zealand reveal the skeletal plan of stem penguins. Journal of Vertebrate Paleontology 32(2):235-254
Sister taxon: Kairuku waitaki
Type specimen: OU 22094, a partial skeleton. Its type locality is Waipati Creek, which is in a Duntroonian coastal sandstone in the Kokoamu Greensand Formation of New Zealand.
Ecology: ground dwelling carnivore