Full reference: W. D. Pierce. 1945. Two new fossils from the Upper Miocene of the Puente Hills. Bulletin of the Southern California Academy of Sciences 44(1):3-6
Sister taxa: none
Type specimen: Its type locality is Axis Co. well Rowland No. 1, 2105-2127 feet depth, which is in a Tortonian marine shale in the Puente Formation of California.