Abundance: 0.01 to 15.91% (median 0.62%)

Latitudinal range: -34.1° to 60.4°

Habitats: (36) only

Found in 36 samples

Australia: Mammoth Cave (Late Pleistocene)

France: Romain-la-Roche (Middle Pleistocene)

Germany: Untereichen-Altenstadt 565 m (Early Miocene)

Greece: Lapsarna (Early Miocene)

Indonesia: Braholo Cave (Excavation Unit G6, Assemblage 4) (Late Pleistocene), Ngalau Gupin-A (Middle Pleistocene), Ngalau Gupin-B (Middle Pleistocene)

Iraq: Palegawra Cave (Late Pleistocene)

Italy: San Teodoro Cave (Unit B-I) (Late Pleistocene), San Teodoro Cave (Unit B-II) (Late Pleistocene), M013 Terre Rosse (Late Miocene)

Lesotho: Sehonghong (RF) (Late Pleistocene)

Malawi: Malowa (Holocene)

Russia: Geographical Society Cave (Late Pleistocene), Chagyrskaya Cave (Layer 6) (Late Pleistocene), Mezmaiskaya Cave (Layer 2A) (Late Pleistocene), Mezmaiskaya Cave (Layer 2B) (Late Pleistocene), Cheremukhovo-1 (Bed 2) (Holocene), Cheremukhovo-1 (Bed 3) (Holocene), Cheremukhovo-1 (Bed 4) (Holocene), Cheremukhovo-1 (Bed 5) (Holocene), Cheremukhovo-1 (Bed 7) (Holocene), Cheremukhovo-1 (Bed 8) (Pleistocene - Holocene), Khayrgas Cave - Layer 6 (1998-1999 pit) (Late Pleistocene)

Sri Lanka: Fa-Hien Lena Cave (Phase A) (Holocene)

Thailand: Spirit Cave (Layer 3) (Late Pleistocene), Doi Pha Kan (Late Pleistocene)

Ukraine: Grot Skeliastyi (III-3-x) (Late Pleistocene)

United States: Thomas Farm (Unit 15) (Early Miocene), Thomas Farm (Unit 11) (Early Miocene), Thomas Farm (Unit 6-7) (Early Miocene), Thomas Farm (Unit 5) (Early Miocene), Cueva Quebrada (Late Pleistocene), Kathy's Pit (Pleistocene - Holocene), Smith Creek Canyon (Reddish-brown Silt) (Late Pleistocene)

Vietnam: Hiem Cave (Holocene)

See also Chiroptera