Abundance: 2.94 to 23.28% (median 16.18%)

Latitudinal range: 40.2° to 45.2°

Habitats: (4) only

Found in 4 samples

France: Saint-Vallier (LD3) (Late Pliocene)

Greece: Milia (MIL5) (Late Pliocene)

Italy: Vigna Nuova (lower layer) (Early Pleistocene)

Spain: La Puebla de Valverde (Early Pleistocene)

See also Croizetoceros