Ursus arctos (carnivore)

Mass: 156 kg based on Kingsley et al. 1983

Diet: frugivore-carnivore based on Berns and Hensel 1972

Abundance: 0.01 to 36.13% (median 0.81%)

Latitudinal range: 30.3° to 68.8°

Habitats: (53), boreal forest/taiga (1), desert/xeric shrubland (1), montane grassland (4), temperate broadleaf/mixed forest (3), temperate coniferous forest (3), temperate savanna (2), tundra (1)

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view a larger map

Found in 68 samples

Canada: Grassy Bay (Holocene)

China: Suojia Township, Yanchiwan National Nature Reserve, Songhua River (Late Pleistocene), Kanas National Nature Reserve

Croatia: Northern Velebit National Park

France: Les Canalettes (Layer 3) (Late Pleistocene), Les Canalettes (Layer 4) (Late Pleistocene), Coudoulous I (Faunal Unit II) (Middle Pleistocene), Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Romain-la-Roche (Middle Pleistocene), Lazaret (C II supérieur UA 13-17) (Middle Pleistocene), Payre (ensemble G) (Middle Pleistocene), Payre (ensemble F) (Middle Pleistocene), Payre (ensemble D) (Late Pleistocene), Tourville-la-Rivière (level D1) (Middle Pleistocene)

Georgia: Dzudzuana Cave (lower area, Unit B) (Late Pleistocene)

Iran: Lar Protected Area, Wezmeh Cave (Late Pleistocene), Golestan National Park

Israel: Marj Rabba (Holocene)

Italy: Western Trento Province (2012), Caverna degli Orsi (sondage A, Unit 11.2) (Late Pleistocene), Riparo Biarzo (SU 5) (Late Pleistocene), Riparo Biarzo (SU 4 + 3B) (Holocene), Riparo Biarzo (SU 3A) (Holocene), Mondeval de Sora (SU 8) (Holocene), Mondeval de Sora (SU 31) (Holocene), La Sassa Cave (Late Pleistocene), Grotta Cucgiliana (Late Pleistocene)

Lebanon: Moghr el-Ahwal Cave 3 (Phases F & E) (Late Pleistocene)

North Korea: Chongphadae Cave (Late Pleistocene)

Poland: Borsuka Cave (Layer VI) (Late Pleistocene), Borsuka Cave (Layer III) (Holocene)

Portugal: Gruta de Figueira Brava (Area C) (Late Pleistocene)

Romania: Southern Transylvania, Noviodunum (TPF sector, Middle Ages) (Holocene)

Russia: Geographical Society Cave (Late Pleistocene), Chagyrskaya Cave (Layer 5) (Late Pleistocene), Chagyrskaya Cave (Layer 6) (Late Pleistocene), Cheremukhovo-1 (Bed 4) (Holocene), Imanay Cave (Pleistocene Layer 1) (Late Pleistocene), Imanay Cave (Pleistocene Layer 2) (Late Pleistocene), Imanay Cave (Pleistocene Layer 3) (Late Pleistocene), Imanay Cave (Pleistocene Layer 4) (Late Pleistocene), Khayrgas Cave - Layer 6 (1998–1999 pit) (Late Pleistocene), Khayrgas Cave - Layer 7 (1998–1999 pit) (Late Pleistocene), Byzovaya (Late Pleistocene)

Spain: Cueva del Angel (Late Pleistocene), Bolomor Cave (Level IV) (Middle Pleistocene)

Tunisia: Meninx (Holocene)

Turkey: Yaylacik Research Forest, Marmaris, Barhal Stream, Direkli Cave (Holocene)

United Kingdom: Lynford Quarry (Late Pleistocene), Balderton Sand and Gravel (Middle Pleistocene)

United States: Dalton Highway, Welsh Cave (Late Pleistocene), Wewukiyepuh (Holocene), Polecat Creek (alluvial gravel) (Late Pleistocene), Schulze Cave (layer B) (Holocene), Schulze Cave (layer C1) (Late Pleistocene), Molybdenum Ridge (triggered detections), Molybdenum Ridge (time-lapse detections), Black Rapids Training Area (triggered detections)

Size measurements:
♀ body mass105000 kgN = 45Kingsley et al. 1983
♀ body mass198900 kgN = 19Kingsley et al. 1983
♀ body mass91000 kgN = 24Kingsley et al. 1983
♀ body mass162500 kgN = 18Kingsley et al. 1983
♂ body mass190600 kgN = 41Kingsley et al. 1983
♂ body mass258300 kgN = 10Kingsley et al. 1983
♂ body mass190400 kgN = 24Kingsley et al. 1983
♂ body mass222400 kgN = 19Kingsley et al. 1983

See also Ursus, Ursus americanus, Ursus bibersoni, Ursus boeckhi, Ursus deningeri, Ursus dolinensis, Ursus etruscus, Ursus kudarensis, Ursus maritimus, Ursus minimus, Ursus sp., Ursus spelaeus, Ursus thibetanus, Ursus vitabilis