Abundance: 0.12 to 3.53% (median 0.60%)

Latitudinal range: 32.7° to 45.2°

Habitats: (12) only

Found in 12 samples

France: Saint-Vallier (LD3) (Late Pliocene)

Greece: Milia (MIL5) (Late Pliocene)

Israel: Ubeidiya (Layer II-23) (Early Pleistocene)

Italy: Pirro Nord 13 (Early Pleistocene), Poggio Rosso (Early Pleistocene)

Montenegro: Trlica (Bed 10) (Early Pleistocene)

Spain: Casablanca I (Early Pleistocene), Fuente Nueva 3 (layer 5) (Early Pleistocene), Venta Micena 4-I (Early Pleistocene), Venta Micena 4-II (Early Pleistocene), Barranco Léon (Early Pleistocene), La Puebla de Valverde (Early Pleistocene)

See also Ursus, Ursus americanus, Ursus arctos, Ursus bibersoni, Ursus boeckhi, Ursus deningeri, Ursus dolinensis, Ursus kudarensis, Ursus maritimus, Ursus minimus, Ursus sp., Ursus spelaeus, Ursus thibetanus, Ursus vitabilis