Abundance: 0.03 to 35.57% (median 0.86%)
Latitudinal range: 37.7° to 50.9°Habitats: (20), Mediterranean woodland (2), temperate broadleaf/mixed forest (3)
Found in 25 samples
Czech Republic: Merkur−North (Early Miocene)
France: Coudoulous I (Faunal Unit II) (Middle Pleistocene), Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Coudoulous I (Faunal Unit VII) (Middle Pleistocene)
Germany: Echzell (Early Miocene), Hambach 6C (Middle Miocene), Griesbeckerzell 1a (Middle Miocene)
Greece: Lake Prespa National Park, Ano Metochi-2 (Late Miocene), Ano Metochi-3 (Late Miocene)
Italy: Poggi di Prata, Cava Muracci (Late Pleistocene), Monticino Quarry (Late Miocene)
Montenegro: Great Ulcinj Beach
Poland: Zabia Cave (Layer 21) (Early Pleistocene), Zabia Cave (Layer 19) (Early Pleistocene)
Portugal: Gruta da Aroeira (Layer Xc) (Middle Pleistocene)
Romania: Fălciu-Prut 1 (Late Miocene)
Turkey: Madran Mountain
Ukraine: Grot Skeliastyi (III-3B) (Late Pleistocene), Grot Skeliastyi (III-3-x) (Late Pleistocene), Grot Skeliastyi (III-1) (Holocene)
United States: Pine Bush
See also Natrix longivertebrata, Natrix maura, Natrix merkurensis, Natrix natrix, Natrix sansaniensis, Natrix sipedon, Natrix sp., Natrix sp. 1, Natrix sp. 2, Natrix tessellata