Ursus (carnivore)

Abundance: 0.01 to 99.10% (median 1.47%)

Latitudinal range: 10.7° to 72.8°

Habitats: (176), boreal forest/taiga (6), desert/xeric shrubland (2), montane grassland (4), temperate broadleaf/mixed forest (17), temperate coniferous forest (5), temperate savanna (3), tropical/subtropical coniferous forest (2), tropical/subtropical dry broadleaf forest (8), tropical/subtropical moist broadleaf forest (1), tundra (1)

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view a larger map

Found in 225 samples

Afghanistan: Nuristan Province

Algeria: Ain Boucherit (Upper) (Early Pleistocene)

Armenia: Hovk-1 Cave (Unit 4) (Late Pleistocene)

Bhutan: Lamai Goempa Research Preserve

Bulgaria: Redaka II (Late Pleistocene)

Cambodia: Chhep Wildlife Sanctuary, Boh Dambang (Late Pleistocene)

Canada: Lindsay, Canadian National Railway (spring), Canadian National Railway (summer), Canadian National Railway (fall), Charlie Lake Cave (Zone IV) (Holocene), Learmonth Site (Holocene), Peale Point House 2 (1) (Holocene), Keffer (Midden 57, screened) (Holocene), Keffer (Midden 57, floated) (Holocene), DjPp-8 (level B) (Holocene), Grassy Bay (Holocene), Irving-Johnston (Holocene), Caverne de la Mine (upper infill) (Holocene), Winefred Lake, EMEND site (retention level 100%, replicate 1), EMEND site (retention level 100%, replicate 2), EMEND site (retention level 100%, replicate 3), Algar

China: Wanglang National Nature Reserve, Shennongjia National Nature Reserve, Suojia Township, Nangunhe Nature Reserve, Guanyinshan Nature Reserve, Yanchiwan National Nature Reserve, Ma'anshan Cave (upper layer) (Late Pleistocene), Ma'anshan Cave (lower layer) (Late Pleistocene), Songhua River (Late Pleistocene), Tengchong Section, Foping National Nature Reserve (2020), Kanas National Nature Reserve, Fanjingshan National Nature Reserve, Jinsitai Cave (Layer 7) (Late Pleistocene), Taosi (Holocene), Zhoujiazhuang (Holocene), Mocun Cave (Late Pleistocene), Gouwan (Holocene), Tianluoshan (Holocene), Cunkong Cave (Holocene), Lower Pubu Cave (Late Pleistocene), Wuyun Cave (Middle Pleistocene), Mohui Cave (Early Pleistocene), Ganxian Cave (Middle Pleistocene), Bailongdong (Middle Pleistocene), Yanlidong (Late Stage) (Late Pleistocene), Yanlidong (Early Stage) (Middle Pleistocene), Zhoukoudian Loc. 1 (Layer 3, West Section) (Middle Pleistocene)

Croatia: Northern Velebit National Park, Mujina Pecina (Level B) (Late Pleistocene), Mujina Pecina (Level D2) (Late Pleistocene)

Denmark: Mullerup (waste) (Holocene)

France: Grotte du Bison (Level I) (Late Pleistocene), Mas d'Azil (Late Pleistocene), Les Auzières 2 (Late Pleistocene), Les Canalettes (Layer 2) (Late Pleistocene), Les Canalettes (Layer 3) (Late Pleistocene), Les Canalettes (Layer 4) (Late Pleistocene), Noisetier Cave (Late Pleistocene), Coudoulous I (Faunal Unit II) (Middle Pleistocene), Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Caune de l'Arago (Unit D) (Middle Pleistocene), Saint-Vallier (LD3) (Late Pliocene), Vergisson II (Late Pleistocene), Romain-la-Roche (Middle Pleistocene), Lazaret (C II supérieur UA 13-17) (Middle Pleistocene), Payre (ensemble G) (Middle Pleistocene), Payre (ensemble F) (Middle Pleistocene), Payre (ensemble D) (Late Pleistocene), Orgnac 3 (level 1) (Middle Pleistocene), Tourville-la-Rivière (level D1) (Middle Pleistocene), Soucy 1 (Middle Pleistocene)

Georgia: Ortvale Klde (late Middle Paleolithic) (Late Pleistocene), Ortvale Klde (early Upper Paleolithic) (Late Pleistocene), Dzudzuana Cave (lower area, Unit B) (Late Pleistocene), Kvabebi (Late Pliocene)

Germany: Untermassfeld (Early Pleistocene)

Greece: Milia (MIL5) (Late Pliocene)

India: Dachigam National Park

Iran: Lar Protected Area, Wezmeh Cave (Late Pleistocene), Golestan National Park

Israel: Marj Rabba (Holocene), Ubeidiya (Layer II-23) (Early Pleistocene)

Italy: Western Trento Province (2012), Caverna degli Orsi (sondage A, Unit 10) (Late Pleistocene), Caverna degli Orsi (sondage A, Unit 11.1) (Late Pleistocene), Caverna degli Orsi (sondage A, Unit 11.2) (Late Pleistocene), Riparo Biarzo (SU 5) (Late Pleistocene), Riparo Biarzo (SU 4 + 3B) (Holocene), Riparo Biarzo (SU 3A) (Holocene), Mondeval de Sora (SU 8) (Holocene), Mondeval de Sora (SU 31) (Holocene), Canale delle Acque Alte (Layer C2) (Late Pleistocene), La Sassa Cave (Late Pleistocene), Pirro Nord 13 (Early Pleistocene), Poggetti Vecchi (Middle Pleistocene), Collecurti bonebed (Early Pleistocene), Isernia La Pineta (3 colluvio level) (Middle Pleistocene), Frantoio (Arda River) (Early Pleistocene), Grotta Cucgiliana (Late Pleistocene), Poggio Rosso (Early Pleistocene)

Japan: Torihama (Holocene)

Laos: Tam Ngu Hao 2 (Middle Pleistocene)

Lebanon: Moghr el-Ahwal Cave 3 (Phases F & E) (Late Pleistocene)

Mexico: Ajos, Madera West, Schroeder Site (Structure 1) (Holocene), El Zurdo (Holocene)

Montenegro: Trlica (Bed 10) (Early Pleistocene), Trlica (Bed 5) (Middle Pleistocene)

Morocco: Grotte des Rhinocéros (lower set) (Middle Pleistocene), Grotte des Rhinocéros (upper set) (Middle Pleistocene), Thomas Quarry Hominid Cave (Middle Pleistocene)

Myanmar: Hukaung Valley Wildlife Sanctuary (Outside the Core), Kweenkoh, Mae Nyaw Kee South

North Korea: Chongphadae Cave (Late Pleistocene)

Poland: Borsuka Cave (Layer VI) (Late Pleistocene), Raj Cave (layer 4) (Late Pleistocene), Raj Cave (layer 6) (Late Pleistocene), Borsuka Cave (Layer III) (Holocene), Zabia Cave (Layer 17) (Early Pleistocene)

Portugal: Gruta de Figueira Brava (Area C) (Late Pleistocene)

Romania: Southern Transylvania, Noviodunum (TPF sector, Middle Ages) (Holocene)

Russia: Geographical Society Cave (Late Pleistocene), Zapovednaya (Late Pleistocene), Chagyrskaya Cave (Layer 5) (Late Pleistocene), Chagyrskaya Cave (Layer 6) (Late Pleistocene), Mezmaiskaya Cave (Layer 2A) (Late Pleistocene), Mezmaiskaya Cave (Layer 2B) (Late Pleistocene), Mezmaiskaya Cave (Layer 3) (Late Pleistocene), Cheremukhovo-1 (Bed 4) (Holocene), Imanay Cave (Pleistocene Layer 1) (Late Pleistocene), Imanay Cave (Pleistocene Layer 2) (Late Pleistocene), Imanay Cave (Pleistocene Layer 3) (Late Pleistocene), Imanay Cave (Pleistocene Layer 4) (Late Pleistocene), Khayrgas Cave - Layer 6 (1998-1999 pit) (Late Pleistocene), Khayrgas Cave - Layer 7 (1998–1999 pit) (Late Pleistocene), Byzovaya (Late Pleistocene), Mezmaiskaya Cave (Layer 2) (Late Pleistocene)

Spain: Casablanca I (Early Pleistocene), Huéscar 1 (Middle Pleistocene), Abric Romaní (Level Ja) (Late Pleistocene), Imanolen Arrobia (Late Pleistocene), Cueva del Angel (Late Pleistocene), Fuente Nueva 3 (layer 5) (Early Pleistocene), Bolomor Cave (Level IV) (Middle Pleistocene), Venta Micena 4-I (Early Pleistocene), Venta Micena 4-II (Early Pleistocene), Barranc de la Boella (La Mina 2) (Early Pleistocene), Barranc de la Boella (El Forn 4) (Early Pleistocene), Barranco Léon (Early Pleistocene), Gran Dolina (TD6.3) (Early Pleistocene), La Puebla de Valverde (Early Pleistocene), Alcoy-Mina (Early Pliocene)

Thailand: Khao Yai National Park, Khao Ang Rue Nai Wildlife Sanctuary, Cave of the Monk (Area 1) (Late Pleistocene), Ban Rai rockshelter (Holocene), Tham Lod rockshelter (Unit 1) (Late Pleistocene), Tham Lod rockshelter (Unit 2) (Late Pleistocene), Tham Lod rockshelter (Unit 3) (Late Pleistocene)

Tunisia: Meninx (Holocene)

Turkey: Yaylacik Research Forest, Marmaris, Barhal Stream, Direkli Cave (Holocene)

Ukraine: Prolom II (Late Pleistocene), Medzhibozh 1 (horizon 16a) (Middle Pleistocene)

United Kingdom: Lynford Quarry (Late Pleistocene), Balderton Sand and Gravel (Middle Pleistocene)

United States: Salt Pond Mountain, Boulder County, North Fork of the Cache la Poudre River, North of the Michigamme Reservoir, Dalton Highway, Zesch Cave (Late Pleistocene), Welsh Cave (Late Pleistocene), Vess Cave (Late Pleistocene), Papago Springs Cave (Late Pleistocene), Haystack Cave (lower levels) (Late Pleistocene), Harrodsburg Crevice (Late Pleistocene), Ripplemead (Late Pleistocene), Bat Cave (Late Pleistocene), Angel Site (Subdivision W10D) (Holocene), Wewukiyepuh (Holocene), Fort Center Mound A (Holocene), McLelland (Holocene), Croley-Evans (Holocene), Harris Creek (Holocene), Garden Creek Mound No. 2 (Holocene), Hobson Site (Holocene), Hale Farm (Holocene), Dead Indian Creek (Holocene), Prison Farm (Holocene), Pine Hill (Holocene), Otter Creek No. 2 (Zone 3) (Holocene), Cole Gravel Pit (Holocene), Sachsen Cave Shelter (Holocene), Robinson Cave (Late Pleistocene), Polecat Creek (alluvial gravel) (Late Pleistocene), Schulze Cave (layer B) (Holocene), Schulze Cave (layer C1) (Late Pleistocene), Schulze Cave (layer C2) (Late Pleistocene), Natural Chimneys (Late Pleistocene), Flagg Swamp Rockshelter (Archaic) (Holocene), Molybdenum Ridge (triggered detections), Molybdenum Ridge (time-lapse detections), Black Rapids Training Area (triggered detections), Bull Neck Swamp Research Forest, Smithsonian Conservation Biology Institute (logs, feature), West of US 287, Cumberland Cave (Middle Pleistocene)

Vietnam: Duoi U'Oi (Late Pleistocene), Lang Trang (Cave II, breccia 5) (Late Pleistocene), Coc Muoi (Middle Pleistocene)

See also Ursus americanus, Ursus arctos, Ursus bibersoni, Ursus boeckhi, Ursus deningeri, Ursus dolinensis, Ursus etruscus, Ursus kudarensis, Ursus maritimus, Ursus minimus, Ursus sp., Ursus spelaeus, Ursus thibetanus, Ursus vitabilis