Abundance: 0.02 to 40.76% (median 0.97%)
Latitudinal range: -48.4° to 67.8°Habitats: (147), Mediterranean woodland (1), boreal forest/taiga (1), desert/xeric shrubland (4), mangrove (3), montane grassland (4), temperate broadleaf/mixed forest (8), temperate coniferous forest (1), temperate savanna (2), tropical/subtropical coniferous forest (1), tropical/subtropical dry broadleaf forest (24), tropical/subtropical moist broadleaf forest (26), tropical/subtropical savanna (14), tundra (1)
Found in 237 samples
Argentina: Iguazù National Park (on roads), Cueva Tùnel (Holocene)
Bangladesh: Sundarbans East Wildlife Sanctuary
Belize: Extinction Cave (Late Pleistocene)
Bhutan: Lamai Goempa Research Preserve
Bolivia: Santa Rosa (control)
Brazil: Uacari Sustainable Development Reserve, Floresta Nacional Amapâ (dry season), Floresta Nacional Amapâ (wet season), Parque Estadual do Turvo (trail), Reserva Natural Vale (year 1), Reserva Natural Vale (year 2), Reserva Natural Vale (year 3), Reserva Natural Vale (year 4), Iguaçu National Park, Formosa Military Training Area, Terra do Meio Ecological Station II, Cachoeira Seca Indigenous Land I
Bulgaria: Redaka II (Late Pleistocene)
Burkina Faso: Arly National Park (2017)
Cambodia: Phnom Prich Wildlife Sanctuary, Chhep Wildlife Sanctuary, Boh Dambang (Late Pleistocene), Vat Komnou (Holocene)
Canada: Old Crow River Loc. 11A (square S30E120) (Late Pleistocene)
China: Altun Mountains, Gouli Region, Suojia Township, Nangunhe Nature Reserve, Xishuangbanna National Nature Reserve, Yanchiwan National Nature Reserve, Songhua River (Late Pleistocene), Hunchun National Reserve (winter), Hunchun National Reserve (summer), Zongri type-site (Layer 2) (Holocene), Zongri type-site (Layer 3) (Holocene), Zongri type-site (Layer 4) (Holocene), Foping National Nature Reserve (2020), Muzhuzhuliang (Holocene), Huoshiliang (Holocene), Tianluoshan (Holocene), Cunkong Cave (Holocene), Wuyun Cave (Middle Pleistocene), Mohui Cave (Early Pleistocene), Ganxian Cave (Middle Pleistocene), Bailongdong (Middle Pleistocene)
Costa Rica: Pacuare Nature Reserve
Ethiopia: Belete-Gera National Forest Priority Area (natural forest), Belete-Gera National Forest Priority Area (coffee forest)
France: Grotte du Bison (Level I) (Late Pleistocene), Les Canalettes (Layer 3) (Late Pleistocene), Noisetier Cave (Late Pleistocene), Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Coudoulous I (Faunal Unit VII) (Middle Pleistocene), Bois-de-Riquet (Locus 1) (Early Pleistocene), Caune de l'Arago (Unit D) (Middle Pleistocene), Vergisson II (Late Pleistocene), Romain-la-Roche (Middle Pleistocene), Lazaret (C II supérieur UA 13-17) (Middle Pleistocene), Payre (ensemble G) (Middle Pleistocene), Payre (ensemble F) (Middle Pleistocene), Payre (ensemble D) (Late Pleistocene), Les Pradelles (level 9) (Late Pleistocene), Lunel-Viel (level 9) (Middle Pleistocene)
Germany: Untermassfeld (Early Pleistocene)
Ghana: Mole National Park
India: Savandurga State Forest (monsoon), Mudumalai Tiger Reserve, Valparai Plateau, Similipal Tiger Reserve (camera traps), Akole Tahsil, Dachigam National Park, Kuno Wildlife Sanctuary (good habitat area), Kuno Wildlife Sanctuary (poor habitat area), Kotada Bhadli (Holocene), Badmal (Period IB) (Holocene), Cathedral Cave (MIS 5) (Late Pleistocene)
Indonesia: Bukit Barisan Selatan National Park, Acacia mangium plantation, Braholo Cave (Excavation Unit G6, Assemblage 2) (Late Pleistocene), Ngalau Gupin-A (Middle Pleistocene), Ngalau Gupin-B (Middle Pleistocene)
Iran: Lar Protected Area, Wezmeh Cave (Late Pleistocene), Golestan National Park
Israel: Manot Cave (Area D, Biogenic Unit) (Late Pleistocene), Motza (Early PPNB) (Holocene), Tel Ro'im West (Late PPNB) (Holocene), Ubeidiya (Layer II-23) (Early Pleistocene)
Italy: Caverna degli Orsi (sondage A, Unit 11.1) (Late Pleistocene), Caverna degli Orsi (sondage A, Unit 11.2) (Late Pleistocene), La Sassa Cave (Late Pleistocene), Isernia La Pineta (3 colluvio level) (Middle Pleistocene), Grotta Cucgiliana (Late Pleistocene)
Japan: Matsugae (Middle Pleistocene)
Kenya: Enkapune Ya Muto (RBL2.1) (Holocene), Enkapune Ya Muto (RBL2.2) (Holocene), Enkapune Ya Muto (RBL2.3) (Holocene), Enkapune Ya Muto (DBS) (Holocene), Karungu (Late Pleistocene), Kibogo (Late Pleistocene), Lainyamok (Middle Pleistocene)
Kyrgyzstan: SaryChat Ertash Zapovednik (ridge), Jangart Hunting Reserve (ridge)
Lesotho: Sehonghong (RF) (Late Pleistocene), Sehonghong (RBL/CLBRF) (Late Pleistocene), Sehonghong (BAS) (Late Pleistocene), Sehonghong (MOS) (Late Pleistocene)
Malaysia: Merapoh, Jerangau Forest Reserve, North Selangor Peat Swamp Forest, Jenut Bayek, Jenut Wan Bulan and Fruiting Area, Endau Rompin National Park
Mauritania: Dhar Tichitt Site 38 (Holocene)
Mexico: La Reserva Natural Sierra Nanchititla, Estación CientÃfica Las Joyas, Tolistoque (rainy season), Abra-Tanchipa Biosphere Reserve (survey 1), Huasabas-Sahuaripa Region, Ajos, Reserva de la Biosfera La Encrucijada (secas), Reserva de la Biosfera La Encrucijada (lluvias)
Mongolia: Zaraa Uul (Locus 3) (Holocene)
Montenegro: Trlica (Bed 10) (Early Pleistocene)
Morocco: Gazelle Cave (upper level) (Late Pleistocene), Zouhrah Cave (L 1) (Late Pleistocene), Zouhrah Cave (L 2) (Late Pleistocene), Zouhrah Cave (L 3) (Pleistocene - Holocene), Grotte des Rhinocéros (lower set) (Middle Pleistocene), Thomas Quarry Hominid Cave (Middle Pleistocene)
Myanmar: Hukaung Valley Wildlife Sanctuary (Outside the Core), Kweenkoh, Mae Nyaw Kee South
Namibia: Waterberg Plateau Park (surrounding flatlands), Waterberg Escarpment, Waterberg Plateau Park, Namib-Naukluft National Park, Etosha Region
Nepal: Limi Valley
North Korea: Chongphadae Cave (Late Pleistocene)
Pakistan: Khunjerab National Park
Peru: Los Amigos Conservation Concession (2005), Los Amigos Conservation Concession (2006), Ichigkat Muja - Cordillera del Cóndor National Park (2018), Ichigkat Muja - Cordillera del Cóndor National Park (2019)
Poland: Raj Cave (layer 4) (Late Pleistocene), Raj Cave (layer 6) (Late Pleistocene)
Portugal: Gruta de Figueira Brava (Area C) (Late Pleistocene)
Republic of the Congo: Odzala-Kokoua National Park, Forestry Management Unit Ngombé
Russia: Geographical Society Cave (Late Pleistocene), Zapovednaya (Late Pleistocene), Chagyrskaya Cave (Layer 5) (Late Pleistocene), Chagyrskaya Cave (Layer 6) (Late Pleistocene), Kamenka (Late Pleistocene), Mezmaiskaya Cave (Layer 2A) (Late Pleistocene), Mezmaiskaya Cave (Layer 3) (Late Pleistocene), Cheremukhovo-1 (Bed 9) (Late Pleistocene), Cheremukhovo-1 (Bed 10) (Late Pleistocene), Imanay Cave (Pleistocene Layer 1) (Late Pleistocene), Imanay Cave (Pleistocene Layer 2) (Late Pleistocene), Imanay Cave (Pleistocene Layer 3) (Late Pleistocene), Imanay Cave (Pleistocene Layer 4) (Late Pleistocene), Khayrgas Cave - Layer 5 (1998–1999 pit) (Late Pleistocene)
Saudi Arabia: Ti's al Ghadah (Middle Pleistocene)
Senegal: Drambos Cave, Diouboye (Holocene)
South Africa: Groenefontein Nature Reserve (fynbos), Garden Route National Park (Site 1), Between George and Karatara, Sibudu Cave (Pinkish Grey Sand) (Late Pleistocene), Equus Cave (Unit 1A) (Holocene), Equus Cave (Unit 1B) (Late Pleistocene), Goudveld State Forest (survey cycle 1), Goudveld State Forest (survey cycle 2), Boegoeberg 1 (Late Pleistocene), Plovers Lake (Late Pleistocene), Diepkloof Rock Shelter (MSA - Post HP) (Late Pleistocene), Diepkloof Rock Shelter (MSA - Late HP) (Late Pleistocene), Diepkloof Rock Shelter (MSA - Jack) (Late Pleistocene), Diepkloof Rock Shelter (MSA - Mike) (Late Pleistocene), Boomplaas Cave (Member BLD) (Holocene), Boomplaas Cave (Member BLA) (Holocene), Boomplaas Cave (Member YOL) (Late Pleistocene), Boomplaas Cave (Member BP) (Late Pleistocene), Boomplaas Cave (Member OLP) (Late Pleistocene), Boomplaas Cave (Member BOL) (Late Pleistocene), Boomplaas Cave (Member OCH) (Late Pleistocene), Boomplaas Cave (Member LOH) (Late Pleistocene), Equus Cave (Unit 2A) (Late Pleistocene), Equus Cave (Unit 2B) (Late Pleistocene), Cornelia-Uitzoek (1998 - 2010 collection) (Early Pleistocene), Cornelia-Uitzoek (old collection) (Early Pleistocene), Duinefontein 2 Horizon 2 (Middle Pleistocene)
Spain: Jou Puerta (Late Pleistocene), Imanolen Arrobia (Late Pleistocene), Punta Lucero Quarry (Middle Pleistocene), Bolomor Cave (Level IV) (Middle Pleistocene), Barranc de la Boella (La Mina 2) (Early Pleistocene)
Sudan: Khor Shambat 1 (Mesolithic) (Holocene)
Tanzania: Mwanihana, Ruaha National Park (dry season, random placement), Mwanihana Forest (2013), Mlele Forest Reserve, Tli'ika, Kuumbi Cave (Trench 10, Phase 1b) (Holocene), Kuumbi Cave (Trench 10, Phase 3) (Late Pleistocene), Kuumbi Cave (Trench 10, Phase 4) (Late Pleistocene), Ewass Oldupa (Trench 2) (Early Pleistocene), Ewass Oldupa (Trench 3) (Early Pleistocene)
Thailand: Hala-Bala Wildlife Sanctuary, Cave of the Monk (Area 1) (Late Pleistocene), Ban Rai rockshelter (Holocene), Tham Lod rockshelter (Unit 2) (Late Pleistocene), Doi Pha Kan (Late Pleistocene)
Tunisia: Doukanet el Khoutifa (Holocene)
Turkey: Ulucak Höyük (Level V) (Holocene)
Uganda: Nile Delta
Ukraine: Prolom II (Late Pleistocene)
United Kingdom: Balderton Sand and Gravel (Middle Pleistocene)
United States: Laubach Cave (Late Pleistocene), Rancho La Brea (Pit 91) (Late Pleistocene), Harrodsburg Crevice (Late Pleistocene), Schulze Cave (layer B) (Holocene), Schulze Cave (layer C1) (Late Pleistocene), Kincaid Shelter (Late Pleistocene), Cumberland Cave (Middle Pleistocene), Quitaque Creek (Late Pleistocene)
Venezuela: El Breal de Orocual (ORS20) (Late Pleistocene)
Vietnam: Duoi U'Oi (Late Pleistocene), Lang Trang (Cave II, breccia 5) (Late Pleistocene), Lang Trang (eastern galleries) (Late Pleistocene), Coc Muoi (Middle Pleistocene)
Yemen: Hawf
Zimbabwe: Ingwezi Game Management Project
See also Panthera atrox, Panthera gombaszoegensis, Panthera leo, Panthera onca, Panthera pardus, Panthera sp., Panthera spelaea, Panthera tigris, Panthera uncia