Abundance: 0.03 to 27.91% (median 4.55%)

Latitudinal range: 41.1° to 50.6°

Habitats: (10) only

Found in 10 samples

Czech Republic: Mokrá-Western Quarry (Early Miocene)

Germany: Griesbeckerzell 1a (Middle Miocene)

Greece: Ano Metochi-2 (Late Miocene), Ano Metochi-3 (Late Miocene)

Italy: Monticino Quarry (Late Miocene)

Poland: Zabia Cave (Layer 21) (Early Pleistocene)

Romania: Fălciu-Prut 1 (Late Miocene)

Ukraine: Grot Skeliastyi (III-3B) (Late Pleistocene), Grot Skeliastyi (III-3-x) (Late Pleistocene), Grot Skeliastyi (III-1) (Holocene)

See also Natrix, Natrix longivertebrata, Natrix maura, Natrix merkurensis, Natrix natrix, Natrix sansaniensis, Natrix sipedon, Natrix sp. 1, Natrix sp. 2, Natrix tessellata