Abundance: 0.02 to 4.47% (median 0.61%)

Latitudinal range: 37.9° to 51.4°

Habitats: (9) only

Found in 9 samples

Czech Republic: Merkur−North (Early Miocene)

France: Romain-la-Roche (Middle Pleistocene), Lazaret (C II supérieur UA 13-17) (Middle Pleistocene)

Germany: Hambach 6C (Middle Miocene)

Italy: Monte Argentario (Early Pleistocene)

Kazakhstan: Baikadam (Middle Miocene)

Romania: Subpiatra 2/1R (Middle Miocene)

Spain: Puerto de la Cadena (Early Pliocene)

Ukraine: Grot Skeliastyi (III-3-x) (Late Pleistocene)

See also Vipera, Vipera ammodytes, Vipera aspis, Vipera berus, Vipera lebetina, Vipera natiensis, Vipera palaestinae, Vipera s.l., Vipera sp. 2, Vipera sp. European vipers group, Vipera sp. Oriental vipers group, Vipera sp. Vipera aspis complex, Vipera ursinii